花药壁由外表皮层，中间纤维层，内部营养层和绒毡层构成。
The anther is made up of an outer epidermis, a middle fibrous layer, and an inner nutritive layer, the tapetum.
天竺葵的绒毡层属周原质团绒毡层，其原生质体侵入药室的时间发生在小孢子中大液泡开始形成的时期。
The tapetum of P. hortorum belongs to periplasmodial tapetum. The invasion into the locule occurs when microspore form large vacuole in its cytoplasm.
绒毡层的提前解体导致不育系花药的中层细胞不退化，为证明中层为绒毡层供给营养的假说提供了证据。
The middle layer cell won't degenerate when the tapetum disintegrates prematurely, This proves the hypothesis that the middle layer cells offer nutrients for the tapetum.
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