包括锌和铜在内的金属，是动物的食物的组成成分，所以标记它们在化石中的分布和含量，可以让我们了解1亿年前这种动物的饮食习惯。

Since metals, including zinc and copper, form part of many animals' diets, so mapping them in fossils could shed light on what the animals fed on 100 million years ago.

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作为公式的一部分，他们测定了生物标记物——总nnal含量，它能够反应烟草特异致肺癌成分nnk的含量。

As part of the formula, they measured a biological marker, total NNAL, which indicates the amount of exposure to the tobacco-specific lung cancer-causing agent NNK.

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随着QTL遗传力、样本容量和标记信息含量的增大，QTL位置以及效应估计值的准确度和精确度逐步提高。

The accuracy and precision of estimates of QTL position and effects will be enhanced as the increasing of QTL heritability, sample size and polymorphism information content of molecular marker.

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