次年产生的夏孢子堆为主要初次侵染来源。
The urediospores developed by the hyphae next year are the main primary inocula of the disease.
冬孢子堆呈暗褐色或黑色，覆盖在完整的寄生表皮下显得发亮。
Telia are dark brown or black, and shine from the intact host epidermal covering.
用带有冬孢子堆的白头翁病叶挂在樟子松松针上接种，也获得阳性反应。
The infected leaves of Pulsatilla potens with teliospores were handed up onto the needles of Mongolian pine for inoculating, the reaction was also positive.
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