Urcaryotes
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核仁是真核细胞最显著的标志之一，但关于其进化上的起源迄今仍所知甚少，已发现有迹像表明原始性真核生物贾第虫没有核仁。
Nucleolar structure is one of the most obvious symbols. However, little is known about its evolutionary origin. Some previous studies implied that Giardia has no nucleolus.
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