submarginal initials
幼叶中层既可由亚边缘原始细胞或远轴层叶肉的第一个细胞单独产生，也可由它们二者共同产生。
The middle layer of laminae is brought about either by submarginal initials of by the first cell of the abaxial layer of mesophyll alone or by both of them .
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