It is closely related with population genetic structure, environment heterogeneity and song learning.
它不仅与种群的遗传有关，而且与环境的异质性以及鸣声学习有关。
Genetic heterogeneity, combined with phenotypic imprecision and poor marker coverage, has contributed to the difficulty in defining risk variants.
遗传异质性、表型介定的不精确性和标记覆盖的狭窄范围增加了定义风险变异的难度。
TSC displays genetic heterogeneity with two known genes: TSC1 and TSC2, which was cloned in 1997 and 1993 respectively.
该病具有遗传异质性，致病基因包含TSC1基因和TSC2基因，分别于1997年和1993年被克隆出。
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