No animal's behaviour is controlled by moral principles. Generally speaking, they do not rise from instinctive behaviour to rational behaviour.

没有一种动物的行为是依据道德原则进行的。一般来说，动物的行为不能从本能的行为升华为合理的行为。

youdao

All seem to lack that instinctive understanding of social behaviour that to most women comes as naturally as breathing.

她们似乎缺乏社交悟性，而大多数女性对社交的领悟来得像呼吸一样自然。

youdao

The problem that confronted Dr Whiten and Dr Boesch was how to disentangle which of chimpanzees' many behaviour patterns are genetically instinctive, which are learnt by individuals in isolation (and so are not cultural, because not copied from others) and which are culturally transmitted (by animals copying one another).

ECONOMIST: Chimpanzee behaviour: Culture club | The