In a sub-Saharan context the virus is spread through transmission in warthogs and other wild pig species and can be transmitted by a particular type of tick.

在非洲撒哈拉以南地区，病毒通过疣猪和其它野猪等传播，也可通过特定类型的虱子传播。 在家猪中直接传播是一个日益严重的问题。

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The idea that these and some 30 other behaviour patterns are cultural rather than genetic was supported by the fact that particular sub-species (which differ from one another genetically) are not associated with particular behaviours.

ECONOMIST: Chimpanzee behaviour: Culture club | The