shade-tolerant tree species [林] 耐荫树种
Light-loving species are always among the first invaders, while shade-tolerant species appear later in the succession.
在这个演替过程中，喜阳植物总是第一批“侵入者”，而那些喜阴植物紧随其后出现。
Spruce on sandy land is a shade-tolerant species, the height growth of its seedlings being accelerated under shady condition.
沙地云杉为耐阴性树种，在幼年期一定程度的遮阴条件可以促进树高生长。
There are a number of planting techniques including under-planting which yields a multi-story stand quickly, while allowing the establishment of shade-tolerant species.
这里有几种植被栽植技术，如林下栽植，能够很快形成复合林分，同时适宜耐阴物种的生长。
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