化石来自于俄罗斯乌拉尔山脉（Ural Mountains）南部，以及南非的卡鲁盆地（Karoo Basin）。根据化石统计分析，在二叠纪末的这次大灭绝事件中，陆地脊椎动物属一级的灭绝率达到了78%。
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...最大石油公司皇家荷兰壳牌公司颁发了一份技术操作许可证，依据该许可证，壳牌公司将研究确认位于南非中部地卡罗盆地(Karoo Basin)部分地区地油气潜力。81552
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South Africa (April 2011) The government placed an indefinite moratorium on hydraulic fracturing in the development of shale gas in the Karoo basin.
FORBES: In Depth: A 'Golden Age' for Natural Gas?
In South Africa, which may have the largest shale-gas reserves on the continent, the shales in the Karoo basin have attracted the attention of Shell, which is increasingly billing itself as a gas-focused company.
ECONOMIST: Shale gas should make the world a cleaner, safer place
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