热带和亚热带的一个陆生蕨类大属；有时归入水龙骨科。

Large genus of terrestrial ferns of tropics and subtropics; sometimes placed in family polypodiaceae.

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此外，有冷杉、云杉、松、铁杉和桦等属乔木花粉和杜鹃科、唇形科、莎草科灌木和草本植物花粉及凤尾蕨、水龙骨、卷柏等属孢子。

Besides, pollen of Abies, Picea, Pinus, Tsuga, Betula, Ericaceae, Labiatae and Cyperaceae and spores of Pteris, Polypodium and Selaginella were also observed.

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