棱鳀属鱼类最早分化于中新世早期。

The divergence date of Thryssa species presented here is early Miocene.

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Phase One, the million and a half years ago, in the early--in the late--Miocene and early Pleistocene phases, there was shift where people were less likely to be just going out and gathering foods from every place they could find it.

第一阶段,一百五十万年前,在整个中新世以及更新世的早期,情况开始转变,人们不光靠,出门四处搜集食物

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