But, Dr Grant says this is the first convincing possible mechanism for a "pre-earthquake cue" that aquatic, semi-aquatic and burrowing animals might be able to sense and respond to.

但是，格兰特博士说，这是有关“地震前线索”的第一个令人信服的可能机制，水生的、半水生的以及掘穴的动物可能能感觉到这样的线索并且作出反应。

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Small burrowing animals and also areas where an invertebrate called "feather stars" live could be affected, according to the report commissioned by Scottish Natural Heritage (SNH).

BBC: Western Isles sub-sea cable 'could affect' mud life

Given that things can move between strata due to the activities of burrowing animals, a few more examples would certainly add to confidence that he is right about the date.

ECONOMIST: A cave in South Africa may rewrite human history

Bioerosion: process by which animals, through drilling, grazing and burrowing, erode hard substances, for example rocks and coral reefs.

UNESCO: L O S S A R Y