He said although more fishing was taking place nearer penguin colonies it was often happening later in the season when these colonies were empty.

他说，尽管在靠近企鹅聚居地的地方有更多的捕鱼活动，但这通常发生在季节的晚些时候，在那时这些聚居地都空了。

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He said although more fishing was taking place nearer penguin colonies it was often happening later in the season when these colonies were empty.

他说，尽管在靠近企鹅聚居地的地方有更多的捕鱼活动，但这通常发生在季节的晚些时候，在那时这些聚居地都空了。

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