Holmes crossed two compatible races of ceratocystis ulmi, the causal agent of Dutch elm disease.

霍姆斯曾把荷兰榆病害的致病因子，长喙壳的两个亲和小种杂交。

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Holmes crossed two compatible races of ceratocystis ulmi, the causal agent of Dutch elm disease.

霍姆斯曾把荷兰榆病害的致病因子，长喙壳的两个亲和小种杂交。

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