Among the more significant local titles to lose circulation, the bigger falls included the Ayrshire Post, down by 10% to 20, 000.

BBC: Sharp fall in city evening newspaper sales

The Philadelphia Inquirer, a former Knight-Ridder paper that was acquired last year by McClatchy and then sold to a local investor group, eked out an average daily circulation gain of 0.6% to 352, 592, a notable improvement from the 7.6% year-on-year decline it recorded during the previous six-month period.

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Local and regional newspapers have suffered similar patterns of falling circulation as national titles, which have been shedding readers, many of them seeking their news online, where advertising has also been migrating.

BBC: Sharp fall in city evening newspaper sales

Parallel circulation figures for the Press and Journal, the Dundee Courier and local titles will be published next month.

BBC: Scottish newspaper sales plunge

In March 1998 Wasserstein installed Pollak, the 43-year-old former head of circulation for the New York Times, as top dog and charged him to exhort newly acquired local publications to expand their regional franchises.

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