The Lions head Down Under again in 2013 determined to avenge their defeat by the Australians in 2001.
"He's had a sore knee for the last ten days, " Lions head coach Warren Gatland explained in a media conference.
Lions head coach Ian McGeechan pulled no late selection surprises for the match, with Croft to start at blindside flanker.
Saturday's defeat in Pretoria means the Lions head to Johannesburg for the third Test with the hosts holding a decisive 2-0 lead in the series.
McGeechan was appointed as Lions head coach for the fourth time in May, having masterminded their last series win over the Springboks back in 1997.
But Gatland - who is one of Lions head coach Ian McGeechan's assistants to South Africa - restored Halfpenny for the Six Nations opener against Scotland.
Wales coach Warren Gatland, in the Lions coaching backroom staff in South Africa, transformed Roberts from wing to centre and now Lions head coach Ian McGeechan is certainly reaping rewards.
Lewsey believes that consistent club form, under the eye of the Lions head coach Ian McGeechan, the director of rugby at Wasps, will give him the best chance of making the 2009 Lions tour to South Africa.
Lions loose-head prop Gethin Jenkins is still out - despite training on Monday - with a calf injury.
So Paul James continues alongside Lions tight-head Adam Jones and recalled hooker Rees, who has recovered from a serious groin injury, in the Wales front row.
The flanker will be up against utility back Foden, the Harlequins duo of Danny Care and Jordan Turner-Hall, and Leicester flanker Tom Croft, another surprising omission from head coach Ian McGeechan's Lions squad.
Head coach Ian McGeechan will name his Lions squad next month ahead of the 10-week trip to South Africa.
McGeechan has been head coach on four British and Irish Lions tours and was at the helm for Scotland, Northampton and, most recently, Wasps.
Head coach Ian McGeechan, who confirmed the Lions would not be calling up any replacements, said the loss of Jenkins, Jones, O'Driscoll and Roberts had a major bearing on the second Test.
The Lions were already two casualties down by then, tight-head Murray departing with an ankle injury after just seven minutes and Hook following five minutes later after a lengthy period of treatment for a neck problem.
The head coach was a former openside flanker for the British and Irish Lions.
Tight-head prop Vickery is a veteran of two World Cup finals and two Lions tours and will bring invaluable experience.
The Lions lost prop Euan Murray (ankle) and fly-half James Hook (head) inside the first 15 minutes at Nelson Mandela Bay Stadium in Port Elizabeth.
Winston wound up choosing Columbia over Harvard, feeling a kinship with the school's longtime head coach, Bid Goswami, as well as Goswami's assistant, former Lions star Howie Endelman.
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